"iTunes for Windows" and IE are connected, yes. It uses IE to download all the stuff you order from the iTunes store.
And apparently, earlier versions of IE had caps on the download sizes of files you acquired. In IE6, the download size cap was 2 GB. in IE7, it was 4GB. In IE8, the cap has been lifted.
You don't have to use IE as your default browser. I don't. I use Firefox. But iTunes for Windows will use IE, no matter what you do.
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